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Back in 2014 , James Gunn’sGuardians of the Galaxywas expected to be the MCU ’s first box office bomb , but it ended up being one of its big hits . With a offbeat comedic sensibility , the originalGuardiansmovie was hailed as a breath of reinvigorated air . Gunn turned five obscure cosmic antiheroes from the fringes of the Marvel universe into some of the most recognisable and darling icons around the world . A couple of old age later , Gunn reunited the cast and recaptured this magic inGuardians of the Galaxy Vol . 2 .
The movies themselves order among the MCU ’s best , but their villains have been a mixed bag . TheGuardiansfilms have some of the MCU ’s expectant unsound guys , like the symbolic Ego and the astonishingly sympathetic Nebula , but they also have a couple of generic , forgettable one like Ronan the Accuser .
Ronan The Accuser
Lee Pace is a great player , but he ’s squandered with a one - dimensionalvillain role inGuardians of the Galaxy . Ronan the Accuser is one of the most generic supervillains in the entire MCU . Like fellow cookie - cutter baddies Malekith and Aldrich Killian , Ronan speaks in diabolical soundbites about his obscure program for world supremacy .
liken to how amusing and eccentric the ease of the cast is , the by - the - numbers baddie is a vast let - down . Ronan poses a serious threat to the Guardians , but the character himself is n’t particularly memorable .
Ayesha
Ayesha is one of the many villain inGuardians of the Galaxy Vol . 2 . Played by Elizabeth Debicki , Ayesha is the High Priestess of the Sovereign , a genetically engineered race of Au - skinned super - the great unwashed . The Sovereign snipe the Guardians with a remotely aviate space fleet at the get-go of theGuardianssequel and again at the end . This al-Qur’an - end function does n’t give Debicki a lot to do , but Ayesha is still a memorable character .
The physical idol of the self-governing people uproariously highlights the glaring imperfectness of the Guardians . In one of theGuardianssequel ’s many credit rating scene , Ayesha is go through creating Adam Warlock for the exclusive function of destroying the Guardians of the Galaxy , so it does n’t seem like fans have seen the last of her .
Thanos
Thanos would n’t become the de facto villain of the entire MCU until he terrorized Earth ’s Mightiest Heroes on his wayto collect the Infinity Stones . But a few years before that , he had a brief supporting role in the firstGuardiansmovie . As usual , the Mad Titan spends his scenes sitting atop a can , deliver expo and give orders to his supporting villain . After bossing Loki around inThe Avengers , Thanos boss Ronan and Nebula around inGuardians of the Galaxy . This version of Thanos has the same bluish aim from the firstAvengersmovie before his comics - accurate purple skin was bring in inInfinity War . His theatrical role inGuardiansmainly serves to place up his role as the gravid badness of the Infinity Saga .
Thanos ’ scenes inGuardians of the Galaxystop the picture dead to establish connection between this motion-picture show and the wider MCU , but at least the producer in conclusion cast Josh Brolin , who went on to grow the sore Titan into one of the most memorable and iconic villain in movie history with his role inthe third and fourthAvengersfilms .
Ego
Peter Quill finally meet his biologic father , Ego , inGuardians of the Galaxy Vol . 2 – and he turn out to be an evil daredevil . Ego is a living planet whose immense top executive has naturally given him delusions of nobility . He wants to reshape the universe in his image , which tie into a great father - Logos story about Quill realizing he knew his existent father all along .
Yondu might not be Quill ’s biological father – and he might not have a perfect trail phonograph record as a parent – but he always loved Quill like he was his own Word . Ego ’s role as a genocidal madman helps to hammer home this content . His cave full of skulls demonstrates how much unsound Yondu ’s parenting could ’ve been . Kurt Russell brings the same unnerving magic spell to this cosmic lord that he bring to the similarly murderous Stuntman Mike in Quentin Tarantino ’s underrated carsploitation slasherDeath Proof .
Nebula
Although Ego is a lot of sport , the greatest villain in theGuardiansmovies is the antihero who terminate up becoming a Guardian herself . Karen Gillan has brought plenty of depth to Gamora ’s sadistic sister , Nebula . The fiber is introduce as a cold - hearted , part - cybernetic killer , but she turns out to be surprisingly kindly . She ’s frame as the secondary scoundrel of the firstGuardiansfilm , but she ’s much more interesting than the main villain . Unlike Ronan , Nebula has a personal connection to the Guardians ( at least to one of them ) and a relatable motive for require to wipe them out .
Nebula is do-or-die to prove she ’s better than Gamora after being buy by Thanos ’ opprobrious , manipulative parenting proficiency . Across the first and secondGuardiansfilms and later in theAvengersmovies , Nebula charted a compelling repurchase spark . Gillan has taken Nebula on a dear journeying from tormenting the Guardians as an foe to becoming a Guardian herself and embracing her alienated sister ’s love .
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